Cucurbitaceae

Cucurbitaceae; Vremenski raspon: od ranog paleocena do sadašnjosti (62–0 Ma) PreЄ Є O S D C P T J K Pg N
	Hodgsonia muški oblik biljke
Naučna klasifikacija
Carstvo:	Plantae
Kladus:	Tracheophytes
Kladus:	Angiospermae
Kladus:	Eudicotidae
Kladus:	Rosids
Red:	Cucurbitales
Porodica:	Cucurbitaceae; Juss.
Tipski rod
	Cucurbita; L.
Plemena i rodovi
	Pogledajte tekst

Bundeve ili tikve (lat. Cucurbitaceae), porodica je dikotiledonih skrivenosemenica koja sadrži oko 965 vrsta raspoređenih u oko 95 rodova.^[2] Prirodni areal rasprostranjenja obuhvata tropske i suptropske oblasti planete. U ovu porodicu spadaju poznate kulturne biljke poput krastavca, lubenice, tikvi, tikvica, bundeva.

Opis[uredi | uredi izvor]

Biljke ove porodice su puzavice, penjačice ili lijane, najčešće uglastog stabla, sa relativno krupnim listovima prekrivenim oštrim dlakama.^[3] Cvetovi su najčešće žute boje (ponekad crvene), jednopolni. Biljke su dvodome ili ređe jednodome. Plod je bobica, sa čvrstom korom (čime podseća na čauru). Seme je spljošteno.

Klasifikacija[uredi | uredi izvor]

Plemenska klasifikacija[uredi | uredi izvor]

Najnovija klasifikacija Cucurbitaceae razgraničava 15 plemena:^[4]^[5]

Pleme Gomphogyneae Benth. & Hook.f.
- Alsomitra (Blume) Spach (1 sp.)
- Bayabusua (1 sp.)
- Gomphogyne Griff. (2 spp.)
- Gynostemma Blume (10 spp.)
- Hemsleya Cogn. ex F.B.Forbes & Hemsl. (30 spp.)
- Neoalsomitra Hutch. (12 spp.)
Pleme Triceratieae A.Rich.
- Anisosperma Silva Manso (1 sp.)
- Cyclantheropsis Harms (3 spp.)
- Fevillea L. (8 spp.)
- Pteropepon (Cogn.) Cogn. (5 spp.)
- Sicydium Schltdl. (7 spp.)
Pleme Zanonieae Benth. & Hook.f.
- Gerrardanthus Harvey in Hook.f. (3–5 spp.)
- Siolmatra Baill. (1 sp.)
- Xerosicyos Humbert (5 spp.)
- Zanonia L. (1 sp.)
Pleme Actinostemmateae H.Schaef. & S.S.Renner
- Actinostemma Griff. (3 spp.)
Pleme Indofevilleeae H.Schaef. & S.S.Renner
- Indofevillea Chatterjee (1 sp.)
Pleme Thladiantheae H.Schaef. & S.S.Renner
- Baijiania A.M.Lu & J.Q.Li (30 spp.)
- Thladiantha Bunge 1833 (5 spp.)
Pleme Siraitieae H. Schaef. & S.S. Renner
- Siraitia Merr. (3–4 spp.)
Pleme Momordiceae H.Schaef. & S.S.Renner
- Momordica L. (60 spp.)
Pleme Joliffieae Schrad.
- Ampelosicyos Thouars (5 spp.)
- Cogniauxia Baill. (2 spp.)
- Telfairia Hook. (3 spp.)
Pleme Bryonieae Dumort.
- Austrobryonia H.Schaef. (4 spp.)
- Bryonia L. (10 spp.)
- Ecballium A.Rich. (1 sp.)
Pleme Schizopeponeae C.Jeffrey
- Herpetospermum Wall. ex Hook.f. (3 spp.)
- Schizopepon Maxim. (6–8 spp.)
Pleme Sicyoeae Schrad.
- Cyclanthera Schrad. (40 spp.)
- Echinocystis Torr. & A.Gray (1 sp.)
- Echinopepon Naudin (20 spp., including Brandegea Cogn.)
- Frantzia Pittier (5 spp.)
- Hanburia Seem. (7 spp.)
- Hodgsonia Hook.f. & Thomson (2 spp.)
- Linnaeosicyos H.Schaef. & Kocyan (1 sp.)
- Luffa Mill. (5–7 spp.)
- Marah Kellogg (7 spp.)
- Nothoalsomitra Hutch. (1 sp.)
- Sicyos L. (75 spp., including Sechium P.Browne)
- Trichosanthes L. (≤100 spp.)
Pleme Coniandreae Endl.
- Apodanthera Arn. (16 spp.)
- Bambekea Cogn. (1 sp.)
- Ceratosanthes Adans. (4 spp.)
- Corallocarpus Welw. ex Benth. & Hook.f. (17 spp.)
- Cucurbitella Walp. (1 sp.)
- Dendrosicyos Balf.f. (1 sp.)
- Doyerea Grosourdy (1 sp.)
- Eureiandra Hook.f. (8 spp.)
- Gurania (Schltdl.) Cogn. (37 spp.)
- Halosicyos Mart.Crov (1 sp.)
- Helmontia Cogn. (2–4 spp.)
- Ibervillea Greene (9–10 spp.)
- Kedrostis Medik. (28 spp.)
- Melotrianthus M.Crovetto (1–3 spp.)
- Psiguria Neck. ex Arn. (6–12 spp.)
- Seyrigia Keraudren (6 spp.)
- Trochomeriopsis Cogn. (1 sp.)
- Tumamoca Rose (2 spp.)
- Wilbrandia Silva Manso (5 spp.)
Pleme Benincaseae Ser.
- Acanthosicyos Welw. ex Hook.f. (1 sp.)
- Benincasa Savi (2 spp., including Praecitrullus Pangalo)
- Borneosicyos (1–2 spp.)
- Cephalopentandra Chiov. (1 sp.)
- Citrullus Schrad. (4 spp.)
- Coccinia Wight & Arn. (30 spp.)
- Ctenolepis Hook. f. 1867 (3 spp.)
- Cucumis L. (65 spp.)
- Dactyliandra Hook.f. (2 spp.)
- Diplocyclos (Endl.) T.Post & Kuntze (4 spp.)
- Indomelothria (2 spp.)
- Khmeriosicyos (1 sp.)
- Lagenaria Ser. (6 spp.)
- Lemurosicyos Keraudren (1 sp.)
- Melothria L. (12 spp., including M. scabra)
- Muellerargia Cogn. (2 sp.)
- Papuasicyos (8 spp.)
- Peponium Engl. (20 spp.)
- Raphidiocystis Hook.f. (5 spp.)
- Ruthalicia C.Jeffrey (2 spp.)
- Scopellaria W.J.de Wilde & Duyfjes (2 spp.)
- Solena Lour. (3 spp.)
- Trochomeria Hook.f. (8 spp.)
- Zehneria Endl. (ca. 60 spp.)
Pleme Cucurbiteae Ser.
- Abobra Naudin (1 sp.)
- Calycophysum H.Karst. & Triana (5 spp.)
- Cayaponia Silva Manso (50–59 spp., including Selysia Cogn.)
- Cionosicys Griseb. (4–5 spp.)
- Cucurbita L. (15 spp.)
- Penelopeia Urb. (2 spp.)
- Peponopsis Naudin (1 sp.)
- Polyclathra Bertol. (6 spp.)
- Schizocarpum Schrad. (11 spp.)
- Sicana Naudin (4 spp.)
- Tecunumania Standl. & Steyerm. (1 sp.)

Sistematika[uredi | uredi izvor]

Savremena molekularna filogenija predlaže sledeće odnose:^[4]^[6]^[7]^[8]^[9]^[10]

Detaljni kladogram koji prikazuje filogeniju familije Cucurbitaceae

Cucurbitaceae

Gomphogyneae

Alsomitra

Bayabusua

	Hemsleya

	Gomphogyne

	Gynostemma

	Neoalsomitra

Triceratieae

Fevillea

Pteropepon

	Cyclantheropsis

	Sicydium

Zanonieae

Gerrardanthus

	Zanonia

	Siolmatra

Xerosicyos

Actinostemmateae	Actinostemma

Indofevilleeae	Indofevillea

Thladiantheae

	Baijiania

	Thladiantha

Siraitieae	Siraitia

Momordiceae	Momordica

Joliffieae

Cogniauxia

	Telfairia

	Ampelosicyos

Bryonieae

	Ecballium

	Bryonia

	Echinocystis

	Marah

	Echinopepon

	Frantzia

	Cyclanthera

	Hanburia

Sicyos

Linnaeosicyos

	Luffa

	Trichosanthes

Schizopeponeae

	Schizopepon

	Herpetospermum

Coniandreae

	Bambekea

	Eureiandra

Dendrosicyos

	Trochomeriopsis

	Seyrigia

Doyerea

	Helmontia

	Gurania

Melotrianthus

Kedrostis

	Ceratosanthes

	Halosicyos

Apodanthera

	Tumamoca

	Ibervillea

Cucurbiteae

Polyclathra

	Peponopsis

	Cucurbita

	Penelopeia

	Calycophysum

	Sicana

Tecunumania

Schizocarpum

Cionosicys

	Abobra

	Cayaponia

Benincaseae

Zehneria

Citrullus

	Peponium

	Lagenaria

	Solena

	Borneosicyos

Benincasa

Ctenolepis

	Dactyliandra

	Trochomeria

Ruthalicia

	Indomelothria

	Melothria

	Coccinia

	Diplocyclos

	Papuasicyos

	Scopellaria

	Muellerargia

	Cucumis

Reference[uredi | uredi izvor]

^ Angiosperm Phylogeny Group (2009). „An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG III”. Botanical Journal of the Linnean Society. 161 (2): 105—121. doi:10.1111/j.1095-8339.2009.00996.x .
^ Christenhusz, M. J. M. & Byng, J. W. (2016). „The number of known plants species in the world and its annual increase”. Phytotaxa. Magnolia Press. 261 (3): 201—217. doi:10.11646/phytotaxa.261.3.1.
^ Stevens, P. F. (2001 onwards). Cucurbitaceae. Angiosperm Phylogeny Website., Приступљено 14. 08. 2008.
^ ^a ^b Schaefer H, Renner SS (2011). „Phylogenetic relationships in the order Cucurbitales and a new classification of the gourd family (Cucurbitaceae)”. Taxon. 60 (1): 122—138. JSTOR 41059827. doi:10.1002/tax.601011.
^ Schaefer H, Kocyan A, Renner SS (2007). „Phylogenetics of Cucumis (Cucurbitaceae): Cucumber (C. sativus) belongs in an Asian/Australian clade far from melon (C. melo)”. BMC Evolutionary Biology. 7: 58—69. PMC 3225884 . PMID 17425784. doi:10.1186/1471-2148-7-58.
^ Zhang L-B, Simmons MP, Kocyan A, Renner SS (2006). „Phylogeny of the Cucurbitales based on DNA sequences of nine loci from three genomes: Implications for morphological and sexual system evolution” (PDF). Molecular Phylogenetics and Evolution. 39 (2): 305—322. PMID 16293423. doi:10.1016/j.ympev.2005.10.002.
^ Schaefer H, Heibl C, Renner SS (2009). „Gourds afloat: A dated phylogeny reveals an Asian origin of the gourd family (Cucurbitaceae) and numerous oversea dispersal events”. Proc R Soc B. 276 (1658): 843—851. PMC 2664369 . PMID 19033142. doi:10.1098/rspb.2008.1447.
^ de Boer HJ, Schaefer H, Thulin M, Renner SS (2012). „Evolution and loss of long-fringed petals: A case study using a dated phylogeny of the snake gourds, Trichosanthes (Cucurbitaceae)”. BMC Evolutionary Biology. 12: 108. PMC 3502538 . PMID 22759528. doi:10.1186/1471-2148-12-108.
^ Belgrano MJ (2012). Estudio sistemático y biogeográfico del género Apodanthera Arn. (Cucurbitaceae) [Systematic and biogeographic study of the genus Apodanthera Arn. (Cucurbitaceae)] (Ph.D.). Universidad Nacional de La Plata.
^ Renner SS, Schaefer H (2016). „Phylogeny and evolution of the Cucurbitaceae”. Ur.: Grumet R, Katzir N, Garcia-Mas J. Genetics and Genomics of Cucurbitaceae. Plant Genetics and Genomics: Crops and Models. 20. New York, NY: Springer International Publishing. str. 1—11. ISBN 978-3-319-49330-5. doi:10.1007/7397_2016_14.

Literatura[uredi | uredi izvor]

Bates D, Robinson R, Jeffrey C, ur. (1990). Biology and Utilization of the Cucurbitaceae. Cornell University Press. ISBN 978-0-8014-1670-5.
Jeffrey C. (2005). „A new system of Cucurbitaceae”. Bot. Zhurn. 90: 332—335.

Spoljašnje veze[uredi | uredi izvor]

[APGIII2009-1] Angiosperm Phylogeny Group (2009). „An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG III”. Botanical Journal of the Linnean Society. 161 (2): 105—121. doi:10.1111/j.1095-8339.2009.00996.x .

[Christenhusz-Byng2016-2] Christenhusz, M. J. M. & Byng, J. W. (2016). „The number of known plants species in the world and its annual increase”. Phytotaxa. Magnolia Press. 261 (3): 201—217. doi:10.11646/phytotaxa.261.3.1.

[3] Stevens, P. F. (2001 onwards). Cucurbitaceae. Angiosperm Phylogeny Website., Приступљено 14. 08. 2008.

[Schaefer-4] Schaefer H, Renner SS (2011). „Phylogenetic relationships in the order Cucurbitales and a new classification of the gourd family (Cucurbitaceae)”. Taxon. 60 (1): 122—138. JSTOR 41059827. doi:10.1002/tax.601011.

[5] Schaefer H, Kocyan A, Renner SS (2007). „Phylogenetics of Cucumis (Cucurbitaceae): Cucumber (C. sativus) belongs in an Asian/Australian clade far from melon (C. melo)”. BMC Evolutionary Biology. 7: 58—69. PMC 3225884 . PMID 17425784. doi:10.1186/1471-2148-7-58.

[6] Zhang L-B, Simmons MP, Kocyan A, Renner SS (2006). „Phylogeny of the Cucurbitales based on DNA sequences of nine loci from three genomes: Implications for morphological and sexual system evolution” (PDF). Molecular Phylogenetics and Evolution. 39 (2): 305—322. PMID 16293423. doi:10.1016/j.ympev.2005.10.002.

[7] Schaefer H, Heibl C, Renner SS (2009). „Gourds afloat: A dated phylogeny reveals an Asian origin of the gourd family (Cucurbitaceae) and numerous oversea dispersal events”. Proc R Soc B. 276 (1658): 843—851. PMC 2664369 . PMID 19033142. doi:10.1098/rspb.2008.1447.

[8] Boer HJ, Schaefer H, Thulin M, Renner SS (2012). „Evolution and loss of long-fringed petals: A case study using a dated phylogeny of the snake gourds, Trichosanthes (Cucurbitaceae)”. BMC Evolutionary Biology. 12: 108. PMC 3502538 . PMID 22759528. doi:10.1186/1471-2148-12-108.

[9] Belgrano MJ (2012). Estudio sistemático y biogeográfico del género Apodanthera Arn. (Cucurbitaceae) [Systematic and biogeographic study of the genus Apodanthera Arn. (Cucurbitaceae)] (Ph.D.). Universidad Nacional de La Plata.

[10] Renner SS, Schaefer H (2016). „Phylogeny and evolution of the Cucurbitaceae”. Ur.: Grumet R, Katzir N, Garcia-Mas J. Genetics and Genomics of Cucurbitaceae. Plant Genetics and Genomics: Crops and Models. 20. New York, NY: Springer International Publishing. str. 1—11. ISBN 978-3-319-49330-5. doi:10.1007/7397_2016_14.

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