Dictyoptera

Dictyoptera; Vremenski raspon: Carboniferous–Recent 300–0 Ma PreЄ Є O S D C P T J K Pg N
	Temnopteryx sp.
Naučna klasifikacija
Domen:	Eukaryota
Carstvo:	Animalia
Tip:	Arthropoda
Klasa:	Insecta
(nerangirano):	Polyneoptera
Nadred:	Dictyoptera; Latreille, 1829
Redovi
	Blattodea (bubašvabe i termiti); ; Mantodea (bogomoljke); †Umenocoleoidea †Alienopteridae; †Umenocoleidae; ;

Dictyoptera (grč. δίκτυον diktyon „mreža“ i πτερόν pteron „krilo“) je red insekata koji uključuje dva postojeća reda polineopternih insekata: red Bubašvabe (termiti i bubašvabe zajedno)^[1] i red Bogomoljke. Dok svi moderni Dictyoptera imaju kratke jajokrilce, najstariji fosili Dictyoptera imaju duge ovipozitore, slično pripadnicima pravokrilaca.

Klasifikacija i filogenija[uredi | uredi izvor]

Upotreba termina Dictyoptera se menjala tokom godina, i iako je uglavnom bio van upotrebe tokom većeg dela prošlog veka, postaje sve šira upotreba. Obično se smatralo nadredom, sa tri reda termiti, bubašvabe i bogomoljke. U nekim klasifikacijama, međutim, Dictyoptera je prebačen u status reda, a u drugim je red termiti podveden pod bubašvabe dok je Dictyoptera zadržan kao nadred. Bez obzira na to, u svim klasifikacijama sastavne grupe su iste, samo se tretiraju u različitim rangovima. Termiti i bubašvabe su veoma blisko povezani, a ekološki i molekularni podaci ukazuju na vezu sa rodom bubašvaba Cryptocercus.^[2]^[3]

Prema genetskim dokazima, najbliži živi srodnici Dictyoptera su Phasmatodea i zagonetni red Notoptera. Ako se Dictyoptera smatraju nadredom, ovi drugi redovi mogu biti uključeni u njega.^[4]

Evolucioni odnosi zasnovani na Eggleton, Beccaloni & Inward 2007 i modifikovani od strane Evangelista et al 2019, prikazani su na kladogramu:^[5]^[6] Porodice bubašvaba Anaplectidae, Lamproblattidae i Tryonicidae nisu prikazane, ali su smeštene u natporodicu Bubašvaba. Porodice bubašvaba Corydiidae and Ectobiidae su ranije bile poznate kao Polyphagidae and Blattellidae.^[7] Kladogram takođe prikazuje porodicu Alienopteridae (prvobitno pripisanu sopstvenom redu „Alienoptera“) kao sestru Mantodea, ali je kasnije ponovo dodeljena izumrloj superfamiliji Bubašvaba Umenocoleoidea.^[8]

Dictyoptera

	Mantodea (Bubašvabe)

	†Alienopteridae

Blattodea

Blaberoidea

Ectobiidae

Pseudophyllodromiinae

	Blattellinae

	Blaberidae (Džinovske bubašvabe)

Solumblattodea

Corydiodea

	Corydiidae (Peščane bubašvabe itd)

	Nocticolidae (Pećinske bubašvabe itd)

Blattoidea

Blattoidae

	Tryonicidae

	Blattidae (Orijentalni, američki i drugi bubašvabe)

Cryptocercidae (bubašvabe braon kapuljača)

Termitoidae (Termiti)

Mastotermitidae

Euisoptera

	Archotermopsidae

	Hodotermitidae

	Serritermitidae

	Termitidae

Reference[uredi | uredi izvor]

^ Yuan, Mingxuan; Chen, Lei; Yu, Philip S.; Mei, Hong (2014-06-01). „Privacy preserving graph publication in a distributed environment”. World Wide Web. 18 (5): 1481—1517. ISSN 1386-145X. S2CID 10329785. doi:10.1007/s11280-014-0290-4.
^ Lo, Nathan; Tokuda, Gaku; Watanabe, Hirofumi; Rose, Harley; Slaytor, Michael; Maekawa, Kiyoto; Bandi, Claudio; Noda, Hiroaki (2000). „Evidence from multiple gene sequences indicates that termites evolved from wood-feeding cockroaches”. Current Biology. 10 (13): 801—804. PMID 10898984. S2CID 14059547. doi:10.1016/s0960-9822(00)00561-3. .
^ Legendre, Frédéric; Nel, André; Svenson, Gavin J.; Robillard, Tony; Pellens, Roseli; Grandcolas, Philippe (2015). „Phylogeny of Dictyoptera: Dating the Origin of Cockroaches, Praying Mantises and Termites with Molecular Data and Controlled Fossil Evidence”. PLOS ONE. 10 (7): e0130127. Bibcode:2015PLoSO..1030127L. PMC 4511787 . PMID 26200914. doi:10.1371/journal.pone.0130127 . .
^ Cameron, Stephen L.; Barker, Stephen C.; Whiting, Michael F. (2006). „Mitochondrial genomics and the new insect order Mantophasmatodea”. Molecular Phylogenetics and Evolution. 38 (1): 274—279. PMID 16321547. doi:10.1016/j.ympev.2005.09.020. .
^ Eggleton, Paul; Beccaloni, George; Inward, Daegan (2007). „Response to Lo et al”. Biology Letters. 3 (5): 564—565. PMC 2391203 . doi:10.1098/rsbl.2007.0367. .
^ Evangelista, Dominic A.; Wipfler, Benjamin; Béthoux, Olivier; Donath, Alexander; Fujita, Mari; Kohli, Manpreet K.; Legendre, Frédéric; Liu, Shanlin; Machida, Ryuichiro; Misof, Bernhard; Peters, Ralph S.; Podsiadlowski, Lars; Rust, Jes; Schuette, Kai; Tollenaar, Ward; Ware, Jessica L.; Wappler, Torsten; Zhou, Xin; Meusemann, Karen; Simon, Sabrina (2019). „An integrative phylogenomic approach illuminates the evolutionary history of cockroaches and termites (Blattodea)”. Proceedings of the Royal Society B: Biological Sciences. 286 (1895). PMC 6364590 . PMID 30963947. doi:10.1098/rspb.2018.2076. .
^ Beccaloni, George W.; Eggleton, Paul (2011-12-23). „Order Blattodea Brunner von Wattenwyl, 1882. In: Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness”. Zootaxa. 3148 (1): 199. ISSN 1175-5334. doi:10.11646/zootaxa.3148.1.37.
^ Vršanský, Peter; Bechly, Günter; Zhang, Qingqing; Jarzembowski, Edmund A.; Mlynský, Tomáš; Šmídová, Lucia; Barna, Peter; Kúdela, Matúš; Aristov, Danil; Bigalk, Sonia; Krogmann, Lars; Li, Liqin; Zhang, Qi; Zhang, Haichun; Ellenberger, Sieghard; Müller, Patrick; Gröhn, Carsten; Xia, Fangyuan; Ueda, Kyoichiro; Vďačný, Peter; Valaška, Daniel; Vršanská, Lucia; Wang, Bo (2018). „Batesian insect-insect mimicry-related explosive radiation of ancient alienopterid cockroaches”. Biologia. 73 (10): 987—1006. S2CID 52270212. doi:10.2478/s11756-018-0117-3. .

Dodatna literatura[uredi | uredi izvor]

Inward, Daegan; Beccaloni, George; Eggleton, Paul (22. 6. 2007). „Death of an order: a comprehensive molecular phylogenetic study confirms that termites are eusocial cockroaches”. Biology Letters. 3 (3): 331—335. PMC 2464702 . PMID 17412673. doi:10.1098/rsbl.2007.0102. CS1 održavanje: Format datuma (veza)

[1] Yuan, Mingxuan; Chen, Lei; Yu, Philip S.; Mei, Hong (2014-06-01). „Privacy preserving graph publication in a distributed environment”. World Wide Web. 18 (5): 1481—1517. ISSN 1386-145X. S2CID 10329785. doi:10.1007/s11280-014-0290-4.

[2] Lo, Nathan; Tokuda, Gaku; Watanabe, Hirofumi; Rose, Harley; Slaytor, Michael; Maekawa, Kiyoto; Bandi, Claudio; Noda, Hiroaki (2000). „Evidence from multiple gene sequences indicates that termites evolved from wood-feeding cockroaches”. Current Biology. 10 (13): 801—804. PMID 10898984. S2CID 14059547. doi:10.1016/s0960-9822(00)00561-3. .

[3] Legendre, Frédéric; Nel, André; Svenson, Gavin J.; Robillard, Tony; Pellens, Roseli; Grandcolas, Philippe (2015). „Phylogeny of Dictyoptera: Dating the Origin of Cockroaches, Praying Mantises and Termites with Molecular Data and Controlled Fossil Evidence”. PLOS ONE. 10 (7): e0130127. Bibcode:2015PLoSO..1030127L. PMC 4511787 . PMID 26200914. doi:10.1371/journal.pone.0130127 . .

[4] Cameron, Stephen L.; Barker, Stephen C.; Whiting, Michael F. (2006). „Mitochondrial genomics and the new insect order Mantophasmatodea”. Molecular Phylogenetics and Evolution. 38 (1): 274—279. PMID 16321547. doi:10.1016/j.ympev.2005.09.020. .

[5] Eggleton, Paul; Beccaloni, George; Inward, Daegan (2007). „Response to Lo et al”. Biology Letters. 3 (5): 564—565. PMC 2391203 . doi:10.1098/rsbl.2007.0367. .

[6] Evangelista, Dominic A.; Wipfler, Benjamin; Béthoux, Olivier; Donath, Alexander; Fujita, Mari; Kohli, Manpreet K.; Legendre, Frédéric; Liu, Shanlin; Machida, Ryuichiro; Misof, Bernhard; Peters, Ralph S.; Podsiadlowski, Lars; Rust, Jes; Schuette, Kai; Tollenaar, Ward; Ware, Jessica L.; Wappler, Torsten; Zhou, Xin; Meusemann, Karen; Simon, Sabrina (2019). „An integrative phylogenomic approach illuminates the evolutionary history of cockroaches and termites (Blattodea)”. Proceedings of the Royal Society B: Biological Sciences. 286 (1895). PMC 6364590 . PMID 30963947. doi:10.1098/rspb.2018.2076. .

[7] Beccaloni, George W.; Eggleton, Paul (2011-12-23). „Order Blattodea Brunner von Wattenwyl, 1882. In: Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness”. Zootaxa. 3148 (1): 199. ISSN 1175-5334. doi:10.11646/zootaxa.3148.1.37.

[8] Vršanský, Peter; Bechly, Günter; Zhang, Qingqing; Jarzembowski, Edmund A.; Mlynský, Tomáš; Šmídová, Lucia; Barna, Peter; Kúdela, Matúš; Aristov, Danil; Bigalk, Sonia; Krogmann, Lars; Li, Liqin; Zhang, Qi; Zhang, Haichun; Ellenberger, Sieghard; Müller, Patrick; Gröhn, Carsten; Xia, Fangyuan; Ueda, Kyoichiro; Vďačný, Peter; Valaška, Daniel; Vršanská, Lucia; Wang, Bo (2018). „Batesian insect-insect mimicry-related explosive radiation of ancient alienopterid cockroaches”. Biologia. 73 (10): 987—1006. S2CID 52270212. doi:10.2478/s11756-018-0117-3. .

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